diff --git a/etc/feature_keys_gff b/etc/feature_keys_gff
index 9f0b36aeaac521546d92ba1942ebb28b56c137f9..a5bcc0d5e7c3fdc79ffcb50a4e35fb5f12ce651a 100644
--- a/etc/feature_keys_gff
+++ b/etc/feature_keys_gff
@@ -10,7 +10,7 @@ region              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ont
 contig              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score comment
 #supercontig         ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
 #chromosome          ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified note comment description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
-gene                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score private
+gene                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score private comment
 #CDS                 ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified codon_start description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score  results note comment job
 polypeptide         ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref product molecule_type size organism_name strain topology localization gff_source gff_seqname score mass isoelectric charge source private signal_peptide membrane_structure cytoplasmic_polypeptide_region non_cytoplasmic_polypeptide_region transmembrane_polypeptide_region cytoplasm_location transmembrane non_cytoplasm_location GPI_anchor_cleavage_site comment curation translation
 pseudogene          ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score private