diff --git a/etc/feature_keys b/etc/feature_keys
index e4e7c4a0e3b671be0d16bb4ad5392cc33ded36a5..d8569eb3a10d209be45dcced39830127d3d1e38c 100644
--- a/etc/feature_keys
+++ b/etc/feature_keys
@@ -10,59 +10,59 @@
3'UTR allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name trans_splicing
5'UTR allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name trans_splicing
CAAT_signal allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial
-CDS EC_number allele artificial_location citation codon_start db_xref exception experiment function gene gene_synonym inference locus_tag map note number old_locus_tag operon partial product protein_id pseudo ribosomal_slippage standard_name trans_splicing transl_except transl_table translation
-C_region allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
+CDS EC_number allele artificial_location citation codon_start db_xref exception experiment function gene gene_synonym inference locus_tag map note number old_locus_tag operon partial product protein_id pseudogene ribosomal_slippage standard_name trans_splicing transl_except transl_table translation
+C_region allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
D-loop allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial
-D_segment allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
+D_segment allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
GC_signal allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial
-J_segment allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
+J_segment allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
LTR allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name
-N_region allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
+N_region allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
RBS allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name
STS allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name
-S_region allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
+S_region allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
TATA_signal allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial
-V_region allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
-V_segment allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
+V_region allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
+V_segment allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
assembly_gap @gap_type @estimated_length linkage_evidence
attenuator allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag operon partial phenotype
enhancer allele bound_moiety citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name
-exon EC_number allele citation db_xref experiment function gene gene_synonym inference locus_tag map note number old_locus_tag partial product pseudo standard_name
+exon EC_number allele citation db_xref experiment function gene gene_synonym inference locus_tag map note number old_locus_tag partial product pseudogene standard_name
gap @estimated_length experiment inference map note
-gene allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial phenotype product pseudo standard_name trans_splicing
+gene allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial phenotype product pseudogene standard_name trans_splicing
iDNA allele citation db_xref experiment function gene gene_synonym inference locus_tag map note number old_locus_tag partial standard_name
-intron allele citation db_xref experiment function gene gene_synonym inference locus_tag map note number old_locus_tag partial pseudo standard_name
-mRNA allele artificial_location citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudo standard_name trans_splicing
-mat_peptide EC_number allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
-misc_RNA allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudo standard_name trans_splicing
+intron allele citation db_xref experiment function gene gene_synonym inference locus_tag map note number old_locus_tag partial pseudogene standard_name
+mRNA allele artificial_location citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudogene standard_name trans_splicing
+mat_peptide EC_number allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
+misc_RNA allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudogene standard_name trans_splicing
misc_binding @bound_moiety allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial
misc_difference allele citation clone compare db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial phenotype replace standard_name
-misc_feature allele citation db_xref experiment function gene gene_synonym inference locus_tag map note number old_locus_tag partial phenotype product pseudo standard_name
+misc_feature allele citation db_xref experiment function gene gene_synonym inference locus_tag map note number old_locus_tag partial phenotype product pseudogene standard_name
misc_recomb allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name
misc_signal allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial phenotype standard_name
misc_structure allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name
mobile_element @mobile_element_type allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial rpt_family rpt_type standard_name
modified_base @mod_base allele citation db_xref experiment frequency gene gene_synonym inference locus_tag map note old_locus_tag
-ncRNA @ncRNA_class allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudo standard_name trans_splicing
+ncRNA @ncRNA_class allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudogene standard_name trans_splicing
old_sequence allele citation compare db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial replace
-operon @operon allele citation db_xref experiment function inference map note phenotype pseudo standard_name
+operon @operon allele citation db_xref experiment function inference map note phenotype pseudogene standard_name
oriT allele bound_moiety citation db_xref direction experiment gene gene_synonym inference locus_tag map note old_locus_tag rpt_family rpt_type rpt_unit_range rpt_unit_seq standard_name
polyA_signal allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial
polyA_site allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag
precursor_RNA allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product standard_name trans_splicing
prim_transcript allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial standard_name
primer_bind PCR_conditions allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name
-promoter allele bound_moiety citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial phenotype pseudo standard_name
+promoter allele bound_moiety citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial phenotype pseudogene standard_name
protein_bind @bound_moiety allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial standard_name
-rRNA allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudo standard_name
+rRNA allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudogene standard_name
rep_origin allele citation db_xref direction experiment gene gene_synonym inference locus_tag map note old_locus_tag partial standard_name
repeat_region allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial rpt_family rpt_type rpt_unit_range rpt_unit_seq satellite standard_name
-sig_peptide allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
+sig_peptide allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
source @mol_type @organism PCR_primers bio_material cell_line cell_type chromosome citation clone clone_lib collected_by collection_date country cultivar culture_collection db_xref dev_stage ecotype environmental_sample focus frequency germline haplogroup haplotype host identified_by isolate isolation_source lab_host lat_lon macronuclear map mating_type note organelle plasmid pop_variant proviral rearranged segment serotype serovar sex specimen_voucher strain sub_clone sub_species sub_strain tissue_lib tissue_type transgenic variety
stem_loop allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial standard_name
-tRNA allele anticodon citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudo standard_name trans_splicing
+tRNA allele anticodon citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag operon partial product pseudogene standard_name trans_splicing
terminator allele citation db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag operon partial standard_name
-tmRNA allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag product pseudo standard_name tag_peptide
-transit_peptide allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudo standard_name
+tmRNA allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag product pseudogene standard_name tag_peptide
+transit_peptide allele citation db_xref experiment function gene gene_synonym inference locus_tag map note old_locus_tag partial product pseudogene standard_name
unsure allele citation compare db_xref experiment gene gene_synonym inference locus_tag map note old_locus_tag replace
variation allele citation compare db_xref experiment frequency gene gene_synonym inference locus_tag map note old_locus_tag partial phenotype product replace standard_name
diff --git a/etc/options b/etc/options
index 5e82d842d63a03c8cdffd723080f701ef14c077c..c856efe5bf3e99fbde3256b1edc999ba7f10d46c 100644
--- a/etc/options
+++ b/etc/options
@@ -531,6 +531,7 @@ extra_qualifiers = \
previous_systematic_id "text" \
primary_name "text" \
prosite_match "text" \
+ pseudo none \
psu_db_xref "text" \
psu_domain "text" \
reserved_name "text" \
@@ -648,6 +649,7 @@ invisible_qualifiers = \
percent_id \
pfam_match \
prosite_match \
+ pseudo \
psu_domain \
reserved_gene_name \
query_id \
diff --git a/etc/qualifier_types b/etc/qualifier_types
index 521250ef6a94088bee33542f0d3f32ae06bff356..7dfd32a0df52e1b7fd5065068ddb82e48499cecc 100644
--- a/etc/qualifier_types
+++ b/etc/qualifier_types
@@ -90,7 +90,7 @@ pop_variant no "text"
product no "text"
protein_id yes "text"
proviral yes none
-pseudo yes none
+pseudogene yes "list" "processed" "unprocessed" "unitary" "allelic" "unknown"
rearranged yes none
replace yes "text"
ribosomal_slippage yes text