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    #
    # Keys (Column 3: "type") constrained to be either:
    # (a) a term from the "lite" sequence ontology, SOFA;
    # (b) a SOFA accession number (SO:000000). 
    #
    # key               Qualifiers (@=mandatory)
    # =======           ========================
    
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    centromere          ID Name Alias Parent Note Target Gap Derives_from feature_id isObsolete timelastmodified note comment private curation
    
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    region              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
    contig              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score comment
    
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    #supercontig         ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
    #chromosome          ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified note comment description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
    
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    gene                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score private
    
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    #CDS                 ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified codon_start description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score  results note comment job
    
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    polypeptide         ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref product molecule_type size organism_name strain topology localization gff_source gff_seqname score mass isoelectric charge source private signal_peptide membrane_structure cytoplasmic_polypeptide_region non_cytoplasmic_polypeptide_region transmembrane_polypeptide_region cytoplasm_location transmembrane non_cytoplasm_location GPI_anchor_cleavage_site comment curation translation
    pseudogene          ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score private
    
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    pseudogenic_exon    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score private
    pseudogenic_transcript ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score product
    intron              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
    
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    exon                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified codon_start description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
    three_prime_UTR     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified systematic_id note comment private curation
    five_prime_UTR      ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified systematic_id note comment private curation
    transcript          ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified private comment
    primary_transcript  ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation 
    tRNA                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
    rRNA                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
    tmRNA                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
    mRNA                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
    ncRNA               ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
    snRNA               ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
    scRNA               ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
    snoRNA              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
    
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    polypeptide_domain  ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified 
    polypeptide_motif   ID Name Alias Parent Note Target Gap Derives_from feature_id isObsolete timelastmodified
    signal_peptide      ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified 
    match               ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified 
    nucleotide_match    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified gff_source gff_seqname score 
    translated_nucleotide_match    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
    protein_match       ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified cluster gff_source gff_seqname score
    sequence_difference ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
    direct_repeat       ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
    inverted_repeat     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
    dispersed_repeat    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
    repeat_family       ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified gff_source gff_seqname score 
    transposable_element  ID Name Alias Parent Target Gap Derives_from Note Dbxref Ontology_term feature_id isObsolete timelastmodified
    binding_site        ID Name Alias Parent Target Gap Derives_from Note Dbxref Ontology_term feature_id isObsolete timelastmodified
    
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    #minus_10_signal     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial standard_name 
    #minus_35_signal     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial standard_name 
    #three_prime_clip    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial standard_name 
    #five_prime_clip     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial standard_name 
    #CAAT_signal         ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
    #D_loop              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
    #GC_signal           ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
    three_prime_LTR     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial standard_name 
    five_prime_LTR      ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial standard_name 
    N_region        	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation codon codon_start evidence gene label locus_tag map note comment partial product pseudo standard_name 
    #STS             	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial standard_name 
    #TATA_box        	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
    #attenuator      	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment operon partial phenotype 
    #enhancer        	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial standard_name 
    
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    gap             	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified @estimated_length score
    
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    #iDNA            	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment number partial standard_name 
    #mature_peptide  	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified EC_number allele citation codon codon_start evidence function gene label locus_tag map note comment partial product pseudo standard_name 
    #modified_base_site  ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified @mod_base allele citation evidence frequency gene label locus_tag map note comment 
    operon          	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified @operon allele citation evidence function label map note comment phenotype pseudo standard_name 
    polyA_signal_sequence  	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
    polyA_site      		ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment 
    #precursor_RNA   		ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial product standard_name 
    primary_transcript 		ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial standard_name 
    #primer_binding_site     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified PCR_conditions allele citation evidence gene label locus_tag map note comment partial standard_name 
    promoter        		ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial phenotype pseudo standard_name 
    #protein_binding_site    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified @bound_moiety allele citation evidence function gene label locus_tag map note comment partial standard_name 
    origin_of_replication	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation direction evidence gene label locus_tag map note comment partial standard_name 
    repeat_region   	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene insertion_seq label locus_tag map note comment partial rpt_family rpt_type rpt_unit standard_name DNA_transposon 
    repeat_unit     	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial rpt_family rpt_type rpt_unit 
    #satellite_DNA       ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial rpt_family rpt_type rpt_unit standard_name 
    #stem_loop       	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial standard_name 
    #terminator      	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment operon partial standard_name 
    #transit_peptide 	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation codon codon_start evidence function gene label locus_tag map note comment partial product pseudo standard_name
    
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